diff --git a/lld/ELF/Driver.cpp b/lld/ELF/Driver.cpp
--- a/lld/ELF/Driver.cpp
+++ b/lld/ELF/Driver.cpp
@@ -1717,8 +1717,11 @@
   for (InputFile *file : lto->compile()) {
     auto *obj = cast<ObjFile<ELFT>>(file);
     obj->parse(/*ignoreComdats=*/true);
-    for (Symbol *sym : obj->getGlobalSymbols())
-      sym->parseSymbolVersion();
+
+    // Parse '@' in symbol names for non-relocatable output.
+    if (!config->relocatable)
+      for (Symbol *sym : obj->getGlobalSymbols())
+        sym->parseSymbolVersion();
     objectFiles.push_back(file);
   }
 }
diff --git a/lld/test/ELF/lto/version-script2.ll b/lld/test/ELF/lto/version-script2.ll
--- a/lld/test/ELF/lto/version-script2.ll
+++ b/lld/test/ELF/lto/version-script2.ll
@@ -1,18 +1,30 @@
 ; REQUIRES: x86
 
-;; Test we parse symbol versions before LTO, otherwise we may get a symbol
-;; named "foo@@VER1", but not "foo" with the version VER1.
+;; For non-relocatable output, test we parse symbol versions after LTO,
+;; otherwise we may get a symbol named "foo@@VER1", but not "foo" with the
+;; version VER1.
 
 ; RUN: llvm-as %s -o %t.o
 ; RUN: echo "VER1 {};" > %t.script
 ; RUN: ld.lld %t.o -o %t.so -shared --version-script %t.script
-; RUN: llvm-readobj --dyn-syms %t.so | FileCheck %s
+; RUN: llvm-readelf --dyn-syms %t.so | FileCheck %s
 
-; CHECK: Name: foo@@VER1 (
+;; For non-relocatable output, @ in symbol names has no effect on undefined symbols.
+; CHECK:      UND       bar{{$}}
+; CHECK-NEXT: {{[1-9]}} foo@@VER1
+
+;; For relocatable output, @ should be retained in the symbol name.
+;; Don't parse and drop `@VER1`. Also check that --version-script is ignored.
+; RUN: ld.lld %t.o -o %t.ro -r --version-script %t.script
+; RUN: llvm-readelf -s %t.ro | FileCheck %s --check-prefix=RELOCATABLE
+
+; RELOCATABLE:      {{[1-9]}} foo@@VER1
+; RELOCATABLE-NEXT: UND       bar@VER1
 
 target datalayout = "e-m:e-p270:32:32-p271:32:32-p272:64:64-i64:64-f80:128-n8:16:32:64-S128"
 target triple = "x86_64-unknown-linux-gnu"
 
 module asm ".global foo"
-module asm "foo:"
+module asm "foo: call bar"
 module asm ".symver foo,foo@@@VER1"
+module asm ".symver bar,bar@@@VER1"