diff --git a/lld/COFF/InputFiles.cpp b/lld/COFF/InputFiles.cpp
--- a/lld/COFF/InputFiles.cpp
+++ b/lld/COFF/InputFiles.cpp
@@ -688,6 +688,8 @@
       Sym = Symtab->addRegular(this, SymName);
     }
     Symbols.push_back(Sym);
+    if (ObjSym.isUsed())
+      Config->GCRoot.push_back(Sym);
   }
   Directives = Obj->getCOFFLinkerOpts();
 }
diff --git a/lld/test/COFF/used-lto.ll b/lld/test/COFF/used-lto.ll
new file mode 100644
--- /dev/null
+++ b/lld/test/COFF/used-lto.ll
@@ -0,0 +1,15 @@
+; REQUIRES: x86
+; RUN: llvm-as -o %t.obj %s
+; RUN: lld-link -dll -debug -opt:ref -noentry -out:%t.dll %t.obj
+; RUN: llvm-pdbutil dump -publics %t.pdb | FileCheck %s
+
+; CHECK: S_PUB32 {{.*}} `foo`
+
+target datalayout = "e-m:w-i64:64-f80:128-n8:16:32:64-S128"
+target triple = "x86_64-pc-windows-msvc"
+
+@llvm.used = appending global [1 x i8*] [i8* bitcast (void ()* @foo to i8*)], section "llvm.metadata"
+
+define void @foo() {
+  ret void
+}
diff --git a/llvm/include/llvm/LTO/LTO.h b/llvm/include/llvm/LTO/LTO.h
--- a/llvm/include/llvm/LTO/LTO.h
+++ b/llvm/include/llvm/LTO/LTO.h
@@ -139,6 +139,7 @@
     using irsymtab::Symbol::getCOFFWeakExternalFallback;
     using irsymtab::Symbol::getSectionName;
     using irsymtab::Symbol::isExecutable;
+    using irsymtab::Symbol::isUsed;
   };
 
   /// A range over the symbols in this InputFile.