Index: wasm/LTO.cpp
===================================================================
--- wasm/LTO.cpp
+++ wasm/LTO.cpp
@@ -72,10 +72,11 @@
 BitcodeCompiler::~BitcodeCompiler() = default;
 
 static void undefine(Symbol *S) {
-  if (isa<DefinedFunction>(S))
-    replaceSymbol<UndefinedFunction>(S, S->getName(), 0);
+  if (auto F = dyn_cast<DefinedFunction>(S))
+    replaceSymbol<UndefinedFunction>(F, F->getName(), 0, F->getFile(),
+                                     F->FunctionType);
   else if (isa<DefinedData>(S))
-    replaceSymbol<UndefinedData>(S, S->getName(), 0);
+    replaceSymbol<UndefinedData>(S, S->getName(), 0, S->getFile());
   else
     llvm_unreachable("unexpected symbol kind");
 }
Index: wasm/SymbolTable.cpp
===================================================================
--- wasm/SymbolTable.cpp
+++ wasm/SymbolTable.cpp
@@ -201,7 +201,9 @@
 Symbol *SymbolTable::addDefinedFunction(StringRef Name, uint32_t Flags,
                                         InputFile *File,
                                         InputFunction *Function) {
-  LLVM_DEBUG(dbgs() << "addDefinedFunction: " << Name << "\n");
+  LLVM_DEBUG(dbgs() << "addDefinedFunction: " << Name << " ["
+                    << (Function ? toString(Function->Signature) : "none")
+                    << "]\n");
   Symbol *S;
   bool WasInserted;
   std::tie(S, WasInserted) = insert(Name, File);
@@ -214,8 +216,12 @@
   if (Function)
     checkFunctionType(S, File, &Function->Signature);
 
-  if (shouldReplace(S, File, Flags))
-    replaceSymbol<DefinedFunction>(S, Name, Flags, File, Function);
+  if (shouldReplace(S, File, Flags)) {
+    const WasmSignature *OldSig = cast<FunctionSymbol>(S)->FunctionType;
+    auto NewSym = replaceSymbol<DefinedFunction>(S, Name, Flags, File, Function);
+    if (!NewSym->FunctionType)
+      NewSym->FunctionType = OldSig;
+  }
   return S;
 }
 
@@ -263,7 +269,8 @@
 Symbol *SymbolTable::addUndefinedFunction(StringRef Name, uint32_t Flags,
                                           InputFile *File,
                                           const WasmSignature *Sig) {
-  LLVM_DEBUG(dbgs() << "addUndefinedFunction: " << Name << "\n");
+  LLVM_DEBUG(dbgs() << "addUndefinedFunction: " << Name <<
+             " [" << (Sig ? toString(*Sig) : "none") << "]\n");
 
   Symbol *S;
   bool WasInserted;