Index: llvm/lib/LTO/LTO.cpp
===================================================================
--- llvm/lib/LTO/LTO.cpp
+++ llvm/lib/LTO/LTO.cpp
@@ -455,20 +455,29 @@
     SymbolResolution Res = *ResI++;
     addSymbolToGlobalRes(Used, Sym, Res, 0);
 
-    if (Sym.getFlags() & object::BasicSymbolRef::SF_Undefined)
-      continue;
-    if (Res.Prevailing && Sym.isGV()) {
+    if (Sym.isGV()) {
       GlobalValue *GV = Sym.getGV();
-      Keep.push_back(GV);
-      switch (GV->getLinkage()) {
-      default:
-        break;
-      case GlobalValue::LinkOnceAnyLinkage:
-        GV->setLinkage(GlobalValue::WeakAnyLinkage);
-        break;
-      case GlobalValue::LinkOnceODRLinkage:
-        GV->setLinkage(GlobalValue::WeakODRLinkage);
-        break;
+      if (Res.Prevailing) {
+        if (Sym.getFlags() & object::BasicSymbolRef::SF_Undefined)
+          continue;
+        Keep.push_back(GV);
+        switch (GV->getLinkage()) {
+        default:
+          break;
+        case GlobalValue::LinkOnceAnyLinkage:
+          GV->setLinkage(GlobalValue::WeakAnyLinkage);
+          break;
+        case GlobalValue::LinkOnceODRLinkage:
+          GV->setLinkage(GlobalValue::WeakODRLinkage);
+          break;
+        }
+      } else if (GV->hasAvailableExternallyLinkage()) {
+        // We can link available_externally symbols even if they are
+        // non-prevailing.
+        GlobalValue *CombinedGV =
+            RegularLTO.CombinedModule->getNamedValue(GV->getName());
+        if (!CombinedGV || CombinedGV->isDeclaration())
+          Keep.push_back(GV);
       }
     }
     // Common resolution: collect the maximum size/alignment over all commons.
Index: llvm/lib/Linker/IRMover.cpp
===================================================================
--- llvm/lib/Linker/IRMover.cpp
+++ llvm/lib/Linker/IRMover.cpp
@@ -870,9 +870,6 @@
   if (DGV && !DGV->isDeclarationForLinker())
     return false;
 
-  if (SGV.hasAvailableExternallyLinkage())
-    return true;
-
   if (SGV.isDeclaration() || DoneLinkingBodies)
     return false;
 
Index: llvm/lib/Linker/LinkModules.cpp
===================================================================
--- llvm/lib/Linker/LinkModules.cpp
+++ llvm/lib/Linker/LinkModules.cpp
@@ -425,7 +425,8 @@
     return;
 
   // Add these to the internalize list
-  if (!GV.hasLinkOnceLinkage() && !shouldLinkOnlyNeeded())
+  if (!GV.hasLinkOnceLinkage() && !GV.hasAvailableExternallyLinkage() &&
+      !shouldLinkOnlyNeeded())
     return;
 
   if (shouldInternalizeLinkedSymbols())
Index: llvm/test/LTO/Resolution/X86/Inputs/link-odr-availextern-ae.ll
===================================================================
--- /dev/null
+++ llvm/test/LTO/Resolution/X86/Inputs/link-odr-availextern-ae.ll
@@ -0,0 +1,6 @@
+target datalayout = "e-m:e-i64:64-f80:128-n8:16:32:64-S128"
+target triple = "x86_64-unknown-linux-gnu"
+
+define available_externally i32 @f() {
+  ret i32 2
+}
Index: llvm/test/LTO/Resolution/X86/link-odr-availextern.ll
===================================================================
--- /dev/null
+++ llvm/test/LTO/Resolution/X86/link-odr-availextern.ll
@@ -0,0 +1,25 @@
+; Tests for correct behavior for non-prevailing resolutions in cases involving
+; *_odr and available_externally linkages.
+
+; RUN: llvm-as %s -o %t1
+; RUN: llvm-as %S/Inputs/link-odr-availextern-ae.ll -o %t2ae
+
+; RUN: llvm-lto2 -o %t3 %t1 %t2ae -r %t1,f,p -r %t2ae,f, -save-temps
+; RUN: llvm-dis < %t3.0.0.preopt.bc -o - | FileCheck --check-prefix=PREVAILING %s
+
+; RUN: llvm-lto2 -o %t3 %t2ae %t1 -r %t1,f,p -r %t2ae,f, -save-temps
+; RUN: llvm-dis < %t3.0.0.preopt.bc -o - | FileCheck --check-prefix=PREVAILING %s
+
+; RUN: llvm-lto2 -o %t3 %t2ae -r %t2ae,f, -save-temps
+; RUN: llvm-dis < %t3.0.0.preopt.bc -o - | FileCheck --check-prefix=NONPREVAILING %s
+
+target datalayout = "e-m:e-i64:64-f80:128-n8:16:32:64-S128"
+target triple = "x86_64-unknown-linux-gnu"
+
+; PREVAILING: define weak_odr i32 @f()
+; PREVAILING-NEXT: ret i32 1
+; NONPREVAILING: define available_externally i32 @f()
+; NONPREVAILING-NEXT: ret i32 2
+define linkonce_odr i32 @f() {
+  ret i32 1
+}
Index: llvm/test/Linker/available_externally_a.ll
===================================================================
--- llvm/test/Linker/available_externally_a.ll
+++ llvm/test/Linker/available_externally_a.ll
@@ -1,5 +1,7 @@
 ; RUN: llvm-link %s %p/available_externally_b.ll -S -o - | FileCheck %s
+; RUN: llvm-link %s -S -o - | FileCheck --check-prefix=AE-ONLY %s
 
 @foo = available_externally unnamed_addr constant i32 0
 
 ; CHECK: @foo = hidden unnamed_addr constant i32 0
+; AE-ONLY-NOT: @foo